| Gnaphalieae | |
|---|---|
![]()  | |
| Helichrysum basalticum | |
| Scientific classification  | |
| Kingdom: | Plantae | 
| Clade: | Tracheophytes | 
| Clade: | Angiosperms | 
| Clade: | Eudicots | 
| Clade: | Asterids | 
| Order: | Asterales | 
| Family: | Asteraceae | 
| Subfamily: | Asteroideae | 
| Tribe: | Gnaphalieae Cass. ex Lecoq & Juillet  | 
| Genera | |
| 
 See text  | |

The pubescent foliage of Helichrysum orientale
The Gnaphalieae are a tribe of flowering plants in the family Asteraceae. It is most closely related to the tribes Anthemideae, Astereae, and Calenduleae.[1]
Characteristics
This group is most diverse in South America, Southern Africa and Australia. There are only a few genera with species native to temperate regions: Anaphalis, Antennaria, Gamochaeta, Helichrysum, Leontopodium (Edelweiss), Phagnalon, Diaperia, and Pseudognaphalium.[2]
The classification of the tribe into subtribes is unclear, with a number of past classifications not being supported by late 20th-century evidence.[2]
Genera
Gnaphalieae genera recognized by the Global Compositae Database as April 2022:[3][4][5]
- × Filfia Holub
 - Acanthocladium F.Muell.
 - Achyrocline (Less.) DC.
 - Acomis F.Muell.
 - Actinobole Fenzl ex Endl.
 - Alatoseta Compton
 - Ammobium R.Br.
 - Amphiglossa DC.
 - Anaphalioides (Benth.) Kirp.
 - Anaphalis DC.
 - Anaxeton Gaertn.
 - Ancistrocarphus A.Gray
 - Anderbergia B.Nord.
 - Anemocarpa Paul G.Wilson
 - Angianthus J.C.Wendl.
 - Antennaria Gaertn.
 - Apalochlamys Cass.
 - Argentipallium Paul G.Wilson
 - Argyroglottis Turcz.
 - Argyrotegium J.M.Ward & Breitw.
 - Arrowsmithia DC.
 - Asteridea Lindl.
 - Athrixia Ker Gawl.
 - Atrichantha Hilliard & B.L.Burtt
 - Basedowia E.Pritz.
 - Bellida Ewart
 - Belloa J.Rémy
 - Berroa Beauverd
 - Blennospora A.Gray
 - Bombycilaena (DC.) Smoljan.
 - Bryomorphe Harv.
 - Calocephalus R.Br.
 - Calomeria Vent.
 - Calotesta P.O.Karis
 - Cassinia R.Br.
 - Castroviejoa Galbany, L.Sáez & Benedí
 - Catatia Humbert
 - Cephalipterum A.Gray
 - Cephalosorus A.Gray
 - Chamaepus Wagenitz
 - Chevreulia Cass.
 - Chiliocephalum Benth.
 - Chionolaena DC.
 - Chondropyxis D.A.Cooke
 - Chryselium Urtubey & S. E. Freire
 - Chrysocephalum Walp.
 - Chthonocephalus Steetz
 - Cladochaeta DC.
 - Coronidium Paul G. Wilson
 - Craspedia G.Forst.
 - Cremnothamnus Puttock
 - Cuatrecasasiella H.Rob.
 - Dasyanthus Bubani
 - Decazesia F.Muell.
 - Diaperia Nutt.
 - Dicerothamnus Koek.
 - Dielitzia P.S.Short
 - Disparago Gaertn.
 - Dithyrostegia A.Gray
 - Dolichothrix Hilliard & B.L.Burtt
 - Edmondia Cass.
 - Elytropappus Cass.
 - Epitriche Turcz.
 - Eriochlamys Sond. & F.Muell.
 - Erymophyllum Paul G.Wilson
 - Euchiton Cass.
 - Ewartia Beauverd
 - Ewartiothamnus Anderb.
 - Facelis Cass.
 - Feldstonia P.S.Short
 - Filago Loefl.
 - Fitzwillia P.S.Short
 - Fluminaria N.G.Bergh
 - Galeomma Rauschert
 - Gamochaeta Wedd.
 - Gilberta Turcz.
 - Gilruthia Ewart
 - Gnaphaliothamnus Kirp.
 - Gnaphalium L.
 - Gnephosis Cass.
 - Gnomophalium Greuter
 - Gratwickia F.Muell.
 - Haeckeria F.Muell.
 - Haegiela P.S.Short & Paul G.Wilson
 - Haptotrichion Paul G.Wilson
 - Helichrysopsis Kirp.
 - Helichrysum Mill.
 - Hesperevax (A.Gray) A.Gray
 - Humeocline Anderb.
 - Hyalochlamys A.Gray
 - Hyalosperma Steetz
 - Hydroidea P.O.Karis
 - Ifloga Cass.
 - Ixiolaena Benth.
 - Ixodia R.Br.
 - Jalcophila M.O.Dillon & Sagást.
 - Lachnospermum Willd.
 - Langebergia Anderb.
 - Laphangium (Hilliard & B.L.Burtt) Tzvelev
 - Lasiopogon Cass.
 - Lawrencella Lindl.
 - Leiocarpa Paul G.Wilson
 - Lemooria P.S.Short
 - Leontopodium (Pers.) R.Br. ex Cass.
 - Lepidostephium Oliv.
 - Leptorhynchos Less.
 - Leucochrysum (A.Cunn. ex DC.) Paul G.Wilson
 - Leucogenes Beauverd
 - Leucophyta R.Br.
 - Leysera L.
 - Logfia Cass.
 - Loricaria Wedd.
 - Lucilia Cass.
 - Metalasia R.Br.
 - Mexerion G.L.Nesom
 - Micropsis DC.
 - Micropus L.
 - Millotia Cass.
 - Mniodes (A.Gray) Benth. & Hook.f.
 - Myriocephalus Benth.
 - Myrovernix Koek.
 - Neotysonia Dalla Torre & Harms
 - Nestlera Spreng.
 - Odixia Orchard
 - Oedera L.
 - Ozothamnus R.Br.
 - Paenula Orchard
 - Parantennaria Beauverd
 - Pentatrichia Klatt
 - Petalacte D.Don
 - Phaenocoma D.Don
 - Phagnalon Cass.
 - Pithocarpa Lindl.
 - Plecostachys Hilliard & B.L.Burtt
 - Podolepis Labill.
 - Podotheca Cass.
 - Pogonolepis Steetz
 - Polycalymma F.Muell. & Sond.
 - Pseudognaphalium Kirp.
 - Psilocarphus Nutt.
 - Pterochaeta Steetz
 - Pterygopappus Hook.f.
 - Pycnosorus Benth.
 - Quinetia Cass.
 - Quinqueremulus Paul G.Wilson
 - Rachelia J.M.Ward & Breitw.
 - Raoulia Hook.f. ex Raoul
 - Raouliopsis S.F.Blake
 - Rhetinocarpha Paul G.Wilson & M.A.Wilson
 - Rhodanthe Lindl.
 - Rhynchopsidium DC.
 - Rutidosis DC.
 - Schoenia Steetz
 - Scyphocoronis A.Gray
 - Seriphium L.
 - Siemssenia Steetz
 - Siloxerus Labill.
 - Sinoleontopodium Y.L.Chen
 - Sondottia P.S.Short
 - Stenocline DC.
 - Stenophalium Anderb.
 - Stoebe L.
 - Stuartina Sond.
 - Stylocline Nutt.
 - Swammerdamia DC.
 - Syncarpha DC.
 - Syncephalum DC.
 - Taplinia Lander
 - Tenrhynea Hilliard & B.L.Burtt
 - Thiseltonia Hemsl.
 - Tietkensia P.S.Short
 - Toxanthes Turcz.
 - Trichanthodium Sond. & F.Muell.
 - Triptilodiscus Turcz.
 - Troglophyton Hilliard & B.L.Burtt
 - Vellereophyton Hilliard & B.L.Burtt
 - Waitzia J.C.Wendl.
 - Xerochrysum Tzvelev
 
Other genera
- Callilepis (may not belong in this tribe[2])
 - Libinhania[6]
 - Oxylaena
 - Planea
 - Relhania
 
References
- ↑ Panero, JL; VA Funk (2002-12-30). "Toward a phylogenetic subfamilial classification for the Compositae (Asteraceae)" (PDF). Proceedings of the Biological Society of Washington. Biological Society of Washington. 115 (4): 909–922. Retrieved 2010-08-28.
 - 1 2 3 Randall J. Bayer; Christopher F. Puttock; Scot A. Kelchner (2000). "Phylogeny of South African Gnaphalieae (Asteraceae) based on two noncoding chloroplast sequences". American Journal of Botany. 87 (2): 259–272. doi:10.2307/2656914. JSTOR 2656914. PMID 10675314.
 - ↑ "Gnaphalieae Cass. ex Lecoq & Juill". Global Compositae Database. Compositae Working Group (CWG). 2022. Retrieved 2022-04-21.
 - ↑ "Gnaphaliinae Dumort". Global Compositae Database. Compositae Working Group (CWG). 2022. Retrieved 2022-04-21.
 - ↑ "Relhaniinae Less". Global Compositae Database. Compositae Working Group (CWG). 2022. Retrieved 2022-04-21.
 - ↑ Kilian, Norbert; Galbany-Casals, Mercè; Sommerer, Ronny; Oberprieler, Christoph; Smissen, Rob; Miller, Antony; Rabe, Katharina (2017). "Systematics of Libinhania, a new endemic genus of Gnaphalieae (Asteraceae) from the Socotra archipelago (Yemen), inferred from plastid, low-copy nuclear and nuclear ribosomal DNA loci". Botanical Journal of the Linnean Society. 183 (3): 373–412. doi:10.1093/botlinnean/bow013.
 
External links
 Data related to Gnaphalieae at Wikispecies
 Media related to Gnaphalieae at Wikimedia Commons- UniProt. "Tribe Gnaphalieae". Retrieved 2008-05-16.
 
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